"If a male enters a tunnel that is already occupied by a rival, they fight by locking horns and try to push each other out."

　　“如果一只雄蜣螂進(jìn)入一個已經(jīng)被對手占據(jù)的洞穴，它們就會用觸角抗衡，相互搏斗，努力把對方擠出去。”

　　Knell added that some male dung beetles are smaller and weaker, but do not have to fight for female attention due to their "substantially bigger testicles".

　　克奈爾說，有一些雄蜣螂體積較小，力量較弱，但它們不需要通過搏斗來吸引雌蜣螂的注意，因?yàn)樗鼈兊摹安G丸相當(dāng)大”。

　　"This suggests they sneak behind the back of the other male, waiting until he's looking the other wayfor a chance to mate with the female," he said.

　　他說：“它們會躲在另一只雄蜣螂的背后，趁其不備時與雌蜣螂進(jìn)行交配。”